Several bacteria, yeast and fungi selectively isolated from paper-mill waste-water grew on veratryl alcohol, a key intermediate of lignin metabolism. Penicillium simplicissimum oxidized veratryl alcohol via a NAD(P)+-dependent veratryl alcohol dehydrogenase to veratraldehyde, which was further oxidized to veratric acid in a NAD(P)+-dependent reaction. Veratric-acid-grown cells contained NAD(P)H-dependent O-demethylase activity for veratrate, vanillate and isovanillate. Protocatechuate was cleaved by a protocatechuate 3,4-dioxygenase.
de Jong, E., Beuling, E. E., van der Zwan, R. P., & de Bont, J. A. M. (1990). Degradation of veratryl alcohol by Penicillium simplicissimum. Applied Microbiology and Biotechnology, 34(3), 420-425. https://doi.org/10.1007/BF00170072